A Sourdough bread is made without commercial yeast. Instead, bakers use a starter, a mixture of flour and water that is left to ferment. Wild yeasts and lactic acid bacteria, which live on grain and in the surrounding air, multiply in the mixture and make it bubble. A well kept starter can be used for years, and some bakers say that theirs has been passed down through generations.
B The two groups of microbes play different roles. The yeasts produce carbon dioxide, which makes the dough rise. The bacteria produce acids, which give the bread its slightly sour taste and also help it to stay fresh for longer. A cooler dough tends to produce more acid, while a warmer one favours the yeasts, so bakers can change the flavour of a loaf by changing the temperature of the kitchen.
C Slow fermentation also changes the dough itself. During the long rise, some of the starch and protein in the flour is broken down. Some researchers suggest that this may make the bread easier to digest for certain people, but the evidence is limited. Sourdough made from wheat is not safe for people with coeliac disease, because the gluten is not removed.
D Keeping a starter alive is a matter of routine. It must be fed regularly with fresh flour and water, and part of it is thrown away each time so that it does not grow too large. A starter that smells strongly of nail polish remover is usually hungry. A large survey of home starters found that the place a starter came from explained very little about the microbes in it, and that the flour and the baker's habits were more important.